Skip to Main Content

Apply Now to the University of Wyoming apply now
Levy Lab - Universityof Wyoming
Contact Us

Daniel Levy


Molecular Biology

1000 E. University Ave.

Laramie, WY 82071



Office Location: SIB 2232

Lab Location: SIB 2316

Levy Lab



My NCBI Bibliography


Mishra, S., Levy, D.L. (2022). Nuclear F-actin and lamin A antagonistically modulate nuclear shape. Journal of Cell Science, 135(13):jcs259692. Highlighted on the cover of the July 2022 issue of Journal of Cell Science.

Wesley, C.C., Levy, D.L. (2022). Differentiation-dependent changes in lamin B1 dynamics and lamin B receptor localization. In revision at Molecular Biology of the Cell.

Schibler, A., Jevtic, P., Pegoraro, G., Levy, D.L.*, Misteli, T.* (2022). Identification of epigenetic modulators as determinants of nuclear size and shape. BioRxiv Preprint May 29, 2022. *Corresponding authors

Chen, P., Levy, D.L. (2022). Regulation of organelle size and organization during development. Seminars in Cell and Developmental Biology, S1084-9521(22)00030-1. doi: 10.1016/j.semcdb.2022.02.002.

Mishra, S., Levy, D.L. (2022). Visualizing nuclear pore complexes in Xenopus egg extracts. Methods in Molecular Biology, 2502:395-405.

Jevtic, P., Elliott, K.W., Watkins, S.E., Sreter, J.A., Jovic, K., Lehner, I.B., Baures, P.W., Tsavalas, J.G., Levy, D.L.*, Varga, K.* (2022). An insect antifreeze protein from Anatolica polita enhances the cryoprotection of Xenopus laevis eggs and embryos. Journal of Experimental Biology, 225(4):jeb243662. doi: 10.1242/jeb.243662. *Corresponding authors.


Vukovic, L.D., Chen, P., Mishra, S., White, K.H, Gigley, J.P., Levy, D.L. (2021). Nuclear Transport Factor 2 (NTF2) suppresses WM983B metastatic melanoma by modifying cell migration, metastasis, and gene expression. Scientific Reports, 11:23586.

Gatlin, J.C., Levy, D.L. (2021). Concepts: Organelle Scaling. Encyclopedia of Biochemistry 3rd Edition, 5:107-112, Elsevier Inc. doi:10.1016/B978-0-12-819460-7.00304-2.


Mukherjee, R.N., Sallé, J., Dmitrieff, S., Nelson, K.M., Oakey, J. Minc, N.*, Levy, D.L.* (2020). The perinuclear ER scales nuclear size independently of cell size in early embryos. Developmental Cell, 54(3):395-409. *Corresponding authors. Highlighted as a “Preview”: Chen, H., Good, M.C. (2020). Nuclear sizER in early development. Developmental Cell, 54:297-298. Faculty Opinions (F1000 Prime) recommendations: and

Wesley, C.C., Mishra, S., Levy, D.L. (2020). Organelle size scaling over embryonic development. WIREs Developmental Biology, 9(5):e376. doi: 10.1002/wdev.376.

Levy, D.L. (2020). Cell biology: Tubulin contributes to spindle size scaling. Current Biology, 30(11):R637-R639.

Hatch, E.M., Levy, D.L. (2020). ASCB Annual Meeting Highlight: Nucleus Structure and Dynamics. Molecular Biology of the Cell, 31(6):397.


Chen, P., Tomschik, M., Nelson, K.M., Oakey, J., Gatlin, J.C., Levy, D.L. (2019). Nucleoplasmin is a limiting component in the scaling of nuclear size with cytoplasmic volume. Journal of Cell Biology, 218(12):4063-4078.

Jevtic, P., Schibler, A.C., Wesley, C.C., Pegoraro, G., Misteli, T.*, Levy, D.L.* (2019). The nucleoporin ELYS regulates nuclear size by controlling NPC number and nuclear import capacity. EMBO Reports, e47283 (10.15252/embr.201847283). *Corresponding authors.

Mukherjee, R.N., Levy, D.L. (2019). Rtn4a promotes exocytosis in mammalian cells. Molecular Biology of the Cell, 30(18):2349-2357.

Jevtic, P., Mukherjee, R.N., Chen, P., Levy, D.L. (2019). Altering the levels of nuclear import factors in early Xenopus laevis embryos affects later development. PLoS One, 14(4):e0215740.


Dilsaver, M.R., Chen, P., Thompson, T.A., Reusser, T., Mukherjee, R.N., Oakey, J., Levy, D.L. (2018). Emerin induces nuclear breakage in Xenopus extract and early embryos. Molecular Biology of the Cell, 29(26):3155-3167.

Milunovic-Jevtic, A., Jevtic, P., Levy, D.L., Gatlin, J.C. (2018). In vivo mitotic spindle scaling can be modulated by changing the levels of a single protein: the microtubule polymerase XMAP215. Molecular Biology of the Cell, 29(11):1311-1317.

Chen, P., Levy, D.L. (2018). Nucleus assembly and import in Xenopus laevis egg extract. Cold Spring Harb Protoc. doi:10.1101/pdb.prot097196.

Jevtic, P., Levy, D.L. (2018). Elucidating nuclear size control in the Xenopus model system. Veterinarski Glasnik, 72(1):1-13.


Edens, L.J., Dilsaver, M.R., Levy, D.L. (2017). PKC-mediated phosphorylation of nuclear lamins at a single serine residue regulates interphase nuclear size in Xenopus and mammalian cells. Molecular Biology of the Cell, 28(10):1389-1399.

Jevtic, P., Levy, D.L. (2017). Both nuclear size and DNA amount contribute to midblastula transition timing in Xenopus laevis. Scientific Reports, 7(1):7908.

Nelson, K., Debroy, D., McBride, R., Levy, D., Gatlin, J., Li, D., Oakey, J. (2017). Hydrogel-based Janus structures for asymmetrical aster nucleation. ISA Biomedical Sciences Instrumentation, 512:193.

Jevtic, P., Levy, D.L. (2017). More cytoplasm, more problems. Developmental Cell, 41(3):221-223.


Vukovic, L.D., Jevtic, P., Zhang, Z., Stohr, B.A., Levy, D.L. (2016). Nuclear size is sensitive to NTF2 protein levels in a manner dependent on Ran binding. Journal of Cell Science, 129:1115-1127.

Edens, L.J., Levy, D.L. (2016). A cell-free assay using Xenopus laevis embryo extracts to study mechanisms of nuclear size regulation. Journal of Visualized Experiments, 114:e54173, doi:10.3791/54173.

Vukovic, L.D., Jevtic, P., Edens, L.J., Levy, D.L. (2016). New insights into mechanisms and functions of nuclear size regulation. International Review of Cell and Molecular Biology, 322:1-59.

Mukherjee, R.N., Chen, P., Levy, D.L. (2016). Recent advances in understanding nuclear size and shape. Nucleus, 7(2):167-186.

Jevtic, P., Milunovic-Jevtic, A., Dilsaver, M.R., Gatlin, J.C., Levy, D.L. (2016). Use of Xenopus cell-free extracts to study size regulation of subcellular structures. International Journal of Developmental Biology, 60:277-288.


Jevtic, P., Levy, D.L. (2015). Nuclear size scaling during Xenopus early development contributes to midblastula transition timing. Current Biology, 25:45-52. Highlighted as a UW press release and in the UW Branding Iron.

Jevtic, P., Edens, L.J., Li, X., Nguyen, T., Chen, P., Levy, D.L. (2015). Concentration-dependent effects of nuclear lamins on nuclear size in Xenopus and mammalian cells. Journal of Biological Chemistry, 290(46):27557-71. Highlighted by UW Extension and the Ag College.

Levy, D.L., Heald, R. (2015). Biological scaling problems and solutions in Amphibians. In: Heald, Hariharan, and Wake (eds) Size Control in Biology: From Organelles to Organisms, Cold Spring Harbor Perspectives in Biology, Cold Spring Harbor Laboratory Press, 73-88. doi: 10.1101/cshperspect.a019166.


Edens, L.J., Levy, D.L. (2014). cPKC regulates interphase nuclear size during Xenopus development. Journal of Cell Biology, 206(4):473-83. Highlighted as a JCB biobyte podcast and "Research Highlight": Du Toit, A. (2014). Honey, I shrunk the nucleus. Nature Reviews Molecular Cell Biology, 15(10):633.

Jevtic, P., Edens, L.J., Vukovic, L.D., Levy, D.L. (2014). Sizing and shaping the nucleus: mechanisms and significance. Current Opinion in Cell Biology, 28:16-27.

Edens, L.J., Levy, D.L. (2014). Size scaling of subcellular organelles and structures in Xenopus laevis and Xenopus tropicalis. In: Kloc and Kubiak (eds) Xenopus Development, John Wiley & Sons, Inc., Hoboken, New Jersey, 325-345.

Jevtic, P., Levy, D.L. (2014). Mechanisms of nuclear size regulation in model systems and cancer. Advances in Experimental Medicine and Biology, 773:537-69.


Edens, L.J., White, K.H., Jevtic, P., Li, X., Levy, D.L. (2013). Nuclear size regulation: from single cells to development and disease. Trends in Cell Biology, 23(4):151-9. Highlighted on the cover of the April 2013 issue of Trends in Cell Biology.

Williams, T.L., Levy, D.L. (2013). Assaying cooperativity of protein-DNA interactions using agarose gel electrophoresis. Methods in Molecular Biology, 1054:253-65.


Levy, D.L., Heald, R. (2012). Mechanisms of intracellular scaling. Annual Review of Cell and Developmental Biology, 28:113-35.


Levy, D.L., Heald, R. (2010). Nuclear size is regulated by importin alpha and Ntf2 in Xenopus. Cell, 143(2):288-98. Highlighted as a Cell PaperFlick and "News and Views": Cohen-Fix, O. (2010). Cell biology: import and nuclear size. Nature, 468:513-16. Faculty Opinions (F1000 Prime) recommendation:

Contact Us

Daniel Levy


Molecular Biology

1000 E. University Ave.

Laramie, WY 82071



Office Location: SIB 2232

Lab Location: SIB 2316

1000 E. University Ave. Laramie, WY 82071
UW Operators (307) 766-1121 | Contact Us | Download Adobe Reader

Accreditation | Virtual Tour | Emergency Preparedness | Employment at UW | Privacy Policy | Harassment & Discrimination | Accessibility Accessibility information icon