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Department of Botany|College of Arts and Sciences

Jake Goheen

Assistant Professor

Specialization: Conservation Biology and Community Ecology

Office: Biological Sciences Bldg, 422
Phone: 307-766-55473


BSc Wildlife Biology Kansas State, 1998
MSc Wildlife Science, Purdue University, 202
Ph.D. Biology, University of New Mexico, 2006


Conservation Biology

Research Emphasis

My research interests lie at the nexus of conservation biology and community ecology. Currently, I am devoted to two research programs: 1) examining how the loss of megaherbivores and other wildlife affects rangeland dynamics in savanna ecosystems through the Ungulate Herbivory Under Rainfall Uncertainty (UHURU); and 2) investigating the interplay between community saturation and human disturbances across wildlife assemblages.

Current Research Projects

Landscape change and herbivore extinction in rangeland ecosystems
Plant defense and trait-mediated community assembly in central Kenya
Herbivore coexistence and tree encroachment in central Kenya
Conservation and demography of the globally-endangered hirola antelope
Climate change, predation, and moose population declines in Wyoming

Selected Publications


Goheen, J.R. and T.M. Palmer. Defensive plant-ants stabilize megaherbivore-driven landscape change in an African savanna. Current Biology 20:1768-1772.

Goheen, J.R., T.M. Palmer, F. Keesing, C. Riginos, and T.P. Young. Large herbivores facilitate savanna tree establishment through diverse and indirect pathways. Journal of Animal Ecology 79:372-382.

Palmer, T.M., D.F. Doak, M.L. Stanton, J.L. Bronstein, E.T. Kiers, T.P. Young, J.R. Goheen, and R.M. Pringle. Synergy of multiple partners, including freeloaders, increases host fitness in a multispecies mutualism. 2010. Proceedings of the National Academy of Sciences 107:17234-17239.

Thibault, K.M., S.K.M. Ernest, E.P. White, J.H. Brown, and J.R. Goheen. Long-term insights into consumer-resource dynamics in desert rodents: effects of community composition across guilds. Journal of Mammalogy 91:787-797.


Drever, M.C., J.R. Goheen, and K. Martin. Species-energy theory, pulsed resources, and the regulation of species richness: avian community dynamics during a mountain pine beetle outbreak. Ecology 90:1095-1105.


Ernest, S.K.M, J.H. Brown, K.M. Thibault, E.P. White, and J.R. Goheen. Zero-sum, the niche, and metacommunities: long-term dynamics of community assembly. American Naturalist 172:E257-E269.

Palmer, T.M., M.L. Stanton, T.P. Young, J.R. Goheen, R.M. Pringle, and R. Karban. Putting ant-acacia mutualisms to the fire: Response. Science 319:1759-1761.

Palmer, T.M., M.L. Stanton, T.P. Young, J.R. Goheen, R.M. Pringle, and R. Karban. Breakdown of an ant-plant mutualism follows the loss of large herbivores from an African savanna. Science 319:192-195.


Goheen, J.R., T.P. Young, F. Keesing, and T.M. Palmer. Consequences of herbivory by native ungulates for the reproduction of a savanna tree. Journal of Ecology 95:129-138.

Lusk, J.J., R.K. Swihart, and J.R. Goheen. Correlates of interspecific synchrony and interannual variation in seed production by deciduous trees. Forest Ecology and Management 242:656-670.

Poe, S., J.R. Goheen, and E.P. Hulebak. Convergent exaptation and adaptation in solitary island lizards. Proceedings of the Royal Society Series B 274:2231-2237.

Swihart, R.K., J.R. Goheen, S.A. Schnelker, and C.E Rizkalla. Testing the generality of patch and landscape-level predictors of tree squirrel occurrence at a regional scale. Journal of Mammalogy 88:564-572.


Goheen, J.R., E.P. White, S.K.M. Ernest, and J.H. Brown. Intra-guild compensation regulates species richness in desert rodents: Reply. Ecology 87: 2121-2125.


Goheen, J.R. and R.K. Swihart. Resource selection and predation of North American red squirrels in deciduous forest fragments. Journal of Mammalogy 86:22-28.

Goheen, J.R., E.P. White, S.K.M. Ernest, and J.H. Brown. Intra-guild compensation regulates species richness in desert rodents. Ecology 86:567-573.


Goheen, J.R., F. Keesing, B.F. Allan, D. Ogada, and R.S. Ostfeld. Net effects of large mammals on Acacia seedling survival in an African savanna. Ecology 85:1555-1561.


Goheen, J.R., G.A. Kaufman, and D.W. Kaufman. Effect of body size on reproductive characteristics of the northern grasshopper mouse in north-central Kansas. Southwestern Naturalist 48:427-431.

Goheen, J.R. and R.K. Swihart. Food-hoarding behavior of gray squirrels and North American red squirrels in the central hardwoods region: implications for forest regeneration. Canadian Journal of Zoology 81:1636-1639.

Goheen, J.R., R.K. Swihart, T.M. Gehring, and M.S. Miller. Forces structuring tree squirrel communities in fragmented landscapes: species differences in forest connectivity and carrying capacity. Oikos 102:95-103.

Goheen, J.R., R.K. Swihart, and J.H. Robins. The anatomy of a range expansion: changes in cranial morphology and rates of energy extraction for North American red squirrels from different latitudes. Oikos 102:33-44.

DeWoody, Y.D., R.K. Swihart, B.A. Craig, and J.R. Goheen. Diversity and stability in communities structured by asymmetric resource allocation. American Naturalist 162:514-527.

Swihart, R.K., T.C. Atwood, J.R. Goheen, D. Scheiman, K. Munroe, and T.M. Gehring. Patch occupancy of North American mammals: Is patchiness in the eye of the beholder? Journal of Biogeography 30:1259-1279.


Goheen, J.R., G.A. Kaufman, and D.W. Kaufman. Patterns of reproduction for western harvest mice in north-central Kansas. Prairie Naturalist 34:107-113.


Post, D.M., T.S. Armbrust, E.A. Horne, and J.R. Goheen. Sexual segregation results in differences in content and quality of bison (Bos bison) diets. Journal of Mammalogy 82:407-413.

Schmidt, K.A., J.R. Goheen, and R. Naumann. Incidental nest predation in songbirds: behavioral indicators detect ecological scales and processes. Ecology 82:2937-2947.

Schmidt, K.A., J.R. Goheen, R. Naumann, R.S. Ostfeld, E.M. Schauber, and A.K. Berkowitz. Experimental removal of strong and weak predators: mice and chipmunks preying on songbird nests. Ecology 82:2927-2936.

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