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Department of Zoology and Physiology
Harold L. Bergman, Department Head
428 Biological Sciences Building
Phone: (307) 766-4207, FAX: (307) 766-5625
MERAV BEN-DAVID, B.S. Tel Aviv University 1984; M.S. 1988; Ph.D. University of Alaska 1996; Professor of Zoology and Physiology 2010, 2000.
CRAIG W. BENKMAN, B.A. University of California at Berkeley 1978; M.S. Northern Arizona Sate University 1981; Ph.D. State University of New York at Albany 1985; Robert Berry Professor of Ecology, Professor of Zoology and Physiology 2004.
HAROLD L. BERGMAN, B.A. Eastern Michigan University 1968; M.S. 1971; Ph.D. Michigan State University 1973; Director SENR/IENR 1997; Professor of Zoology and Physiology 1984, 1975.
STEVEN W. BUSKIRK, B.S. University of Redlands 1969; M.S. University of Arizona 1972; Ph.D. University of Alaska 1983; Professor of Zoology and Physiology 1997, 1984.
FRANCIS W. FLYNN, B.A. University of Colorado, Boulder 1973; M.A. Northern Arizona University 1977; Ph.D. Kansas State University 1981; Professor of Zoology and Physiology 2001.
ZOLTAN M. FUZESSERY, B.S. University of California at Santa Barbara 1970; M.S. 1973; Ph.D. University of Illinois 1982; Professor of Zoology and Physiology 2004, 1987.
WILLIAM A. GERN, B.A. Western State College 1971; M.A. 1973; Ph.D. University of Colorado at Boulder 1976; Professor of Zoology and Physiology 1989, 1979.
ROBERT O. HALL JR., B.S. Cornell University 1989; Ph.D. University of Georgia 1996; Professor of Zoology and Physiology 2010, 1998.
HENRY J. HARLOW, B.A. California State University 1966; M.A. 1973; Ph.D. University of Wyoming 1979; Director UW-NPS 1992; Professor of Zoology and Physiology 1996, 1981.
CARLOS MARTINEZ del RIO, B.Sc. Universidad National de Mexico 1984; Ph.D. University of Florida 1990; Professor of Zoology and Physiology 2004, 2000.
DAVID B. MCDONALD, A.B. Harvard College 1973; Ph.D. University of Arizona 1987; Professor of Zoology and Physiology 2010, 1996.
FRANK J. RAHEL, B.A. Kenyon College 1974; M.S. University of Wisconsin 1977; Ph.D. 1982; Professor of Zoology and Physiology 1998, 1985.
ROBERT S. SEVILLE, B.S. San Diego State University 1981; M.S. University of Wyoming 1987; Ph.D. 1992; Professor of Zoology and Physiology 2011, 1995.
DONAL C. SKINNER, B.S. Rhodes University 1987, B.S. University of the Witwatersrand 1988; Ph.D. University of Cambridge 1993; Professor of Zoology and Physiology 2012.
BRUCE W. CULVER, B.S. University of Wyoming 1969; Ph.D. University of Kansas Medical Center 1975; Associate Professor of Pharmacy 1981, 1977, Zoology and Physiology 1984.
QIAN-QUAN SUN, B.Sc. Shandong Normal University 1990; M.S. 1993; Ph.D. St. Andrews University 1998; Associate Professor of Zoology and Physiology 2010, 2004.
CHARLES J. WOODBURY, B.S. Arizona State University 1982; Ph.D. State University of New York at Stony Brook 1989; Associate Professor of Zoology and Physiology 2008, 2003.
MATTHEW D. CARLING, B.S. University of Michigan 1997; M.S. University of Idaho 2002; Ph.D. Louisiana State University 2008; Assistant Professor of Zoology and Physiology 2011.
ANNA D. CHALFOUN, B.A. Smith College 1995; M.S. University of Missouri-Columbia 2000; Ph.D. University of Montana-Missoula 2006; Assistant Assistant Professor of Zoology and Physiology 2011.
BRIAN D. CHERRINGTON, B.A. Washington University 1996; M.S. Colorado State University 2001; Ph.D. 2005; Assistant Professor of Physiology 2011.
MICHAEL E. DILLON, B.S. University of Texas, Austin 1998; Ph.D. University of Washington 2005; Assistant Professor of Zoology and Physiology 2012, 2009.
JACOB R. GOHEEN, B.S. Kansas State University 1998; M.S. Purdue University 2002; Ph.D. University of New Mexico 2006; Assistant Professor of Zoology and Physiology 2012, 2010.
MATTHEW J. KAUFFMAN, B.A. University of Oregon 1992; Ph.D. University of California, Santa Cruz 2003; Assistant Professor of Zoology and Physiology 2006.
AMY M. NAVRATIL, B.S. Colorado State University 1999; Ph.D. 2005; Gardner-Fiske Assistant Professor of Physiology 2011.
JONATHAN F. PRATHER, B.S. University of Virginia 1995; Ph.D. Emory University 2001; Assistant Professor of Zoology and Physiology 2012, 2009.
KARA PRATT, B.A.S. University of of Delaware; Ph.D. Brandeis University 2004; Assistant Professor of Zoology and Physiology 2011.
ANNIKA W. WALTERS, B.A. Princeton University 2002; M.S. Yale University 2006; Ph.D. 2009; Assistant Professor of Zoology and Physiology 2011.
Associate Academic Professional Lecturer
MARGARET J. FLANIGAN, B.Sc. University of Strathclyde 1984; PH.D. University of the Witwatersrand 1993; Academic Professional, Zoology and Physiology 2006, 2003.
Assistant Academic Professional Lecturers
ELIZABETH A. FLAHERTY, B.S. University of Wyoming 2000; M.N.S. Southeast Missouri State University 2002; Ph.D. University of Wyoming 2008; Assistant Academic Professional Lecturer 2011.
CARLY N. JORDAN, B.S. Virginia Polytechnic Institute and State University 2003; M.S. 2005; Ph.D. University of Georgia 2010; Assistant Academic Professional Lecturer 2011.
Senior Academic Professional Research Scientist
ZHAOJIE ZHANG, B.S. Shandong University 1985; M.S. 1988; Ph.D. University of Oklahoma 1999; Director, Microscopy Core Facility, University of Wyoming 2006, 2004; Senior Academic Professional /Research Scientist 2012.
Associate Academic Professional Research Scientist
AMY C. KRIST, B.A. State University of New York at Potsdam 1991; Ph.D. Indiana University 1998; Research Scientist, Zoology and Physiology 2004.
Robert W. Atherton, Robert P. George, Wayne A. Hubert, Robert M. Kitchin, J.A. Lillegraven, Frederick G. Lindzey, James R. Lovvorn, Graham Mitchell, James D. Rose, Joan Smith-Sonneborn
Academic Professional Lecturer Emeritus
Wyoming Cooperative Fish and Wildlife Research Unit
Unit Leader: Matthew W. Kauffman
Assistant Unit Leader for Fisheries: Annika W. Walters
Assistant Unit Leader for Wildlife: Anna D. Chalfoun
Zoology and Physiology
The department of Zoology and Physiology offers a variety of courses in the biological sciences that encompass many aspects of animal form, function, and biology.
Whether you are interested in the intricacies of cell biology or the complexities of ecosystem functioning and whether you want to become a wildlife biologist or a physician, we offer a major that will suit your needs. Students can choose from four undergraduate degrees: biology, physiology, wildlife and fisheries biology and management, or zoology. Course requirements for each degree are detailed on our web site: www.uwyo.edu/zoology.
The Biology major is an interdepartmental program that provides a broad education in biological sciences. It enables students to combine courses in biology, botany, zoology, physiology, and other biological sciences to meet the requirements of the major. On completion of the core requirements for the major, specific courses selected to complete the major may vary according to students' interests and are worked out by consultations between student and adviser. The requirements for a bachelor's degree are outlined in the Biology section of this Bulletin or see the web site at www.uwyo.edu/biology.
Physiology is the study of how animals work: how they breathe, feed, interact with their environment, and carry out many other activities and functions. Physiology is the knowledge that the health sciences are built on and so is especially important for students who may be thinking of becoming medical practitioners, veterinarians or health care professionals. In addition to the University and College of Arts and Sciences requirements, a degree in physiology typically involves the following courses:
Freshman and Sophomore Years
Students take introductory courses in biology, chemistry, physics, and mathematics as these are essential for understanding physiological processes. Students begin the study of physiology with Human Systems Physiology which focuses on how the cardiovascular and respiratory systems function.
Junior and Senior Years
Students take Integrative Physiology, which is concerned with how the body regulates such functions as reproduction or blood glucose concentrations.
Students can specialize in an area of physiology they find particularly interesting. The department has strong expertise in neuroscience physiology, cell physiology, ecological and comparative physiology. For details, visit our web site.
At the end of this program students will have a thorough knowledge of physiology, will be well prepared to enter health sciences or graduate education, and will have a range of skills attractive to employers.
Wildlife and Fisheries Biology and Management Major
Wildlife and Fisheries Biology and Management is a professional degree designed to prepare students for state, federal, and other positions in resource management and conservation biology. The degree provides students with knowledge of the natural world, understanding of processes governing dynamics of wildlife and fish populations, as well as an appreciation of human-mediated effects on wildlife and fish populations. A student graduating with this degree will be familiar with the theory of resource management as well as with methods used to determine population status, habitat quality, and conservation. In Wyoming the abundance of wild animals and pristine habitats provide a unique natural laboratory for studying the responses of wildlife and fish populations to changing climates and habitats.
In addition to the University and College of Arts and Sciences requirements, a degree in wildlife and fisheries biology and management typically involves the following:
Freshman and Sophomore Years
Students take introductory courses in biology, chemistry, physics, and mathematics as these provide essential tools for understanding ideas and processes in wildlife and fisheries biology and management.
Students begin the study of wildlife and fisheries biology and management by taking courses in resource management, natural history of vertebrates, physiology, genetics, and evolution, as these subjects provide the underlying principles of population dynamics and the mechanisms of evolution.
Junior and Senior Years
Students can elect to concentrate in those areas of wildlife and fisheries biology and management they find most interesting and can specialize in a terrestrial or aquatic option. For details of these options see our web site.
A student graduating with a degree in WFBM will have comprehensive knowledge of wildlife and fisheries biology and management, will have earned a degree that is compatible with the requirements for professional certification with the American Fisheries Society or the Wildlife Society, and will have a range of knowledge and skills that are valuable to potential employers.
Zoology is the study of animals: their structure, physiology, development and evolution. One of the enduring fascinations of zoology is that we can learn so much about ourselves and our environment by studying what our fellow creatures do.
In addition to the University and College of Arts and Sciences requirements, a degree in zoology typically involves the following courses:
Students take introductory courses in biology, chemistry, physics, and mathematics as these provide essential tools for understanding zoological ideas and processes.
Students finish any introductory courses they have not completed and begin the study of zoology by taking courses in anatomy, physiology, genetics, ecology, and evolution, as these subjects provide the underlying principles of the mechanisms of evolution, and animal structure, function, and ecology.
Junior and Senior Years
Students take courses in five main areas: invertebrate zoology, ichthyology, herpetology, ornithology, and mammalogy. For details of these and other courses see our Website. At the end of this program students will have a comprehensive knowledge of zoology, will be well prepared for graduate education, and will be equipped to enter any of the many employment opportunities that are available.
Learning Outcomes for Undergraduates
The learning outcomes that direct the teaching of the department's degrees and which we expect our graduates to have acquired are:
- Competence in basic sciences;
- Competence in the content of the specific courses that constitute the principal knowledge of the degree;
- Ability to comprehend, analyze, and interpret biological data where appropriate; and
- Ability to synthesize information from the biological literature, and communicate it effectively in writing or orally.
Minors in zoology, physiology, wildlife fisheries biology management, and neuroscience are offered. Contact the department for further information or see the web site http://www.uwyo.edu/zoology.
The Department of Zoology and Physiology offers programs leading to the master of science and the doctor of philosophy in zoology and physiology. We also participate in graduate programs through the Neuroscience Program and the Program in Ecology.
Program Specific Admission Requirements
Admission is open to all students who meet the minimum requirements set forth in the admissions section of this bulletin.
Research and teaching assistantships are available for graduate students working toward the M.S. or Ph.D. degrees. Applicants can apply for this financial assistance at the time they apply for admission to graduate standing. Applications must be completed by February 15 in order to be considered for the following academic year.
Information on how to apply to the graduate program in the Department of Zoology and Physiology is detailed on our web site. Begin by identifying a faculty member in our department whose research interests are similar to yours. We will only consider an application if a faculty member has indicated a willingness to serve as the student's adviser. After finding a potential adviser, e-mail a completed departmental application form, a copy of your curriculum vitae, copies of college transcripts, recommendation letters and GRE scores to him or her. Our graduate admissions committee will review all applications and make decisions on admission based on the availability of funding and a commitment from a faculty member to serve as the adviser. Students recommended for admission will then be asked to fill out an application to the University of Wyoming and pay a non-refundable application fee.
Consult the web site, www.uwyo.edu/zoology, to find out about faculty research.
Program Specific Degree Requirements
Plan A (thesis)
Includes 26 hours of coursework and 4 hours of thesis research.
Applicants should have at least 20 semester hours of undergraduate work in zoology, physiology, or other areas of the biosciences and have completed introductory courses in mathematics, chemistry, and in at least one other natural or physical science. Early in the second semester the student must file a program of study with the university and have a graduate committee appointed. Plan A candidates shall orally defend the thesis before the graduate committee.
All M.S. candidates will be required to complete credit in two graduate seminars. A student may enroll in more than one of these required seminars during one semester or academic year.
After two semesters in the department, a Plan A master's candidate may request permission from the department's graduate advisory board to proceed directly to the Ph.D. degree; however, such a bypass is granted only by the department head after considering recommendations from the graduate advisory board.
Zoology and physiology may be used as a field by a candidate working for the interdisciplinary master of science in natural science in the College of Arts and Sciences and the College of Education.
Plan B (non-thesis)
Includes 30 hours of coursework.
The program for the Plan B is established by the student and a faculty adviser and must be approved by the department head during the student's second semester or summer session.
The graduate committee will require the candidate to take a written examination. An oral examination may also be required. The final examination is comprehensive, covering all areas of zoology, but emphasizing one major area.
This is a 72 hour program.
A Ph.D. applicant must have 20 hours of undergraduate work in zoology, physiology, or other areas of biology and also have completed substantial undergraduate work in other sciences. Under exceptional circumstances, a student with an undergraduate major in a scientific discipline other than biology may be admitted. After the Ph.D. student has completed two semesters of graduate work, s/he must be approved for continued work toward the doctorate by the graduate advisory board. This board can reconsider a candidate thereafter if it so desires.
A graduate committee shall be appointed for the individual student no later than the third semester. After consultation with the student, this committee will prescribe special requirements (courses, minors, research tools, etc.) that must be fulfilled. At this time, the graduate committee shall consult with the candidate on the proposed research and shall identify the subject matter areas to be included in the preliminary examination. The preliminary examination will consist of a written research proposal, its oral defense, and a written and oral preliminary examination. When training outside zoology and physiology is specified by the committee, certification of satisfactory completion of the requirement will be made by the appropriate department.
In addition to the general university requirements for the Ph.D. degree, the department requires the following:
The coursework program should include work in a discipline outside the department, generally in the sense of a minor, to be identified in consultation with the graduate committee.
The preliminary examination consisting of written and oral portions should be taken no later than midterm of the fourth semester in residence. The graduate committee will certify satisfactory performance for the preliminary examination.
The dissertation must be received by each member of the graduate committee three weeks before the final dissertation seminar. As oral defense of the dissertation, the candidate will deliver a formal 50 minute seminar on original research from the dissertation. The seminar will be followed by an examination by the graduate committee.
Some time during their degree program, all Ph.D. candidates will be required to complete credit in three graduate seminars. A student may enroll in more than one of these required seminars during one semester or academic year.
All candidates for the Ph.D. degree shall be required to teach for one semester during their program.
The dissertation may be written in a format suitable for publication in a journal and the usual extensive literature review, description of study sites, technical details, raw data, supporting figures, charts, and photographs should be included in a well-organized appendix. (See also format requirements by the university.)